CLNEOct 15, 2025

Hierarchical Frequency Tagging Probe (HFTP): A Unified Approach to Investigate Syntactic Structure Representations in Large Language Models and the Human Brain

arXiv:2510.13255v13 citationsh-index: 2Has Code
Originality Incremental advance
AI Analysis

This work addresses the interpretability of LLM improvements for computational linguistics and cognitive neuroscience, though it is incremental as it builds on existing probing and analysis methods.

The researchers tackled the problem of understanding whether large language models (LLMs) process syntax using mechanisms similar to the human brain by introducing the Hierarchical Frequency Tagging Probe (HFTP), which revealed that LLMs like GPT-2 and Llama 3.1 process syntax in analogous layers while the human brain uses distinct cortical regions, with upgraded models showing divergent trends in brain alignment.

Large Language Models (LLMs) demonstrate human-level or even superior language abilities, effectively modeling syntactic structures, yet the specific computational modules responsible remain unclear. A key question is whether LLM behavioral capabilities stem from mechanisms akin to those in the human brain. To address these questions, we introduce the Hierarchical Frequency Tagging Probe (HFTP), a tool that utilizes frequency-domain analysis to identify neuron-wise components of LLMs (e.g., individual Multilayer Perceptron (MLP) neurons) and cortical regions (via intracranial recordings) encoding syntactic structures. Our results show that models such as GPT-2, Gemma, Gemma 2, Llama 2, Llama 3.1, and GLM-4 process syntax in analogous layers, while the human brain relies on distinct cortical regions for different syntactic levels. Representational similarity analysis reveals a stronger alignment between LLM representations and the left hemisphere of the brain (dominant in language processing). Notably, upgraded models exhibit divergent trends: Gemma 2 shows greater brain similarity than Gemma, while Llama 3.1 shows less alignment with the brain compared to Llama 2. These findings offer new insights into the interpretability of LLM behavioral improvements, raising questions about whether these advancements are driven by human-like or non-human-like mechanisms, and establish HFTP as a valuable tool bridging computational linguistics and cognitive neuroscience. This project is available at https://github.com/LilTiger/HFTP.

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